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Extra resources for Aquatic Oligochaete Biology: Proceedings of the 4th International Symposium on Aquatic Oligochaete Biology
Hierarchy and reticulation in systematics. Syst. Zoo!. 28: 624-627. , 1975. Modes of Animal Speciation. Ann. Rev. Eco!. Syst. 6: 339-364. , 1987. Phylogenetic analysis of some Enchytraeidae (Annelida: Oligochaeta): parsimony analysis of structural characters. PhD. dissertation. University of Victoria, Victoria, British Columbia. 392 pp. , 1989. Phylogeny and origins of Enchytraeidae. A. J. Diaz, 1988. Description of Guaranidrilus oregonensis n. sp. (Oligochaeta: Enchytraeidae), the first species of Guaranidrilus from North America, with comments on the genus and other species.
Craggi type H; and Hemienchytraeus brasiliensis. There is no recent phylogenetic analysis to which the clades found here can be compared. Cernosvitov's (1937) classification anticipated some of these but large numbers of species have been described since and the discussion of generic relationships made by Cernosvitov indicated that his subfamilies, especially Achaetinae, and some genera were not conceived as monophylies. Phylogenetic analyses eventually should allow evolutionary investigations of Enchytraeidae to proceed within a theoretically consistent framework and allow the logical reclassification of species in nominal genera such as Marionina.
It seems likely that two different states are confounded as 'secondary glands' (107), one characteristic of the outgroup and early enchytraeids and the other derived from a compact state. Dorsal fusion of pharyngeal glands is characteristic for all enchytraeids, and no clear pattern emerged for the absence of fusion at any or all of the bearing septa. Information on the origin of the dorsal blood vessel (113-115) regarding intraspecific variation and variation at the generic level is probably inaccurate both with regards to specific variation and actual location.
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